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Louisa "Lou" Clark is hired as carer for Will Traynor, a once successful banker and active sportsman now tetraplegic after being hit by a motorcycle. Will's mother hopes Lou's bubbly personality will lift Will's depressed and cynical spirits. Will is initially cold towards Lou. Will's ex-girlfriend Alicia visits and reveals that she is to marry Will's former best friend Rupert. Lou perseveres with Will and, as the two grow close, she learns he is cultured and worldly, in contrast to her simple life spent with her parents or boyfriend Patrick.

Overhearing an argument between Will's parents, Lou learns that Will has conceded six months to them before he will go to Dignitas in Switzerland for assisted suicide, unable as he is to accept a disabled life. Lou takes it upon herself to change his mind by organising trips and adventures to show him that life is still worth living. Patrick's jealousy grows until he and Lou split up.

Will asks Lou to accompany him to Alicia's wedding. They start to fall in love. During a luxurious trip to Mauritius, Will tells Lou he still intends to take assisted suicide, saying he wants her to live a full life instead of half a life with him. Heartbroken, Lou quits as Will's caregiver and refuses contact with him.

Lou's father convinces her to visit Will, but she finds he has already left for Switzerland. She follows him there to be with him in his final moments.

Some weeks after Will's death, sitting in his favourite café in Paris, Lou reads a letter he left for her. In it, he says he has left her enough money to follow her dreams and encourages her to live abundantly.